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Mafia's storyline gameplay consists of driving, mainly easy city cruise between different locations, as well as chases and races; the rest of the game is based on third-person on-foot navigation and shooting - all inter-connected with cutscenes. In addition to the photo-realistic city and a huge countryside, detailed interiors like the city's airport, a museum, a church, a hotel, an abandoned prison, restaurants and Don Salieri's bar are included. Weather changes and day/night cycles are also in use.

Mafia offers an opportunity to drive 51 classic American cars around the city, plus 19 bonus cars (of which 5 are racing models) unlockable after the main mode and the opening of a new game mode. Cars are introduced periodically - in the beginning of the game, early 1920's models drive on the streets of the city, while models from 1930 begin appearing in later game stages.

Police book players for minor offenses such as speeding or running a red light, and car accidents cause physical harm to the driving player. While other forms of transport are available, such as trams and elevated rails, they are only ridable and not drivable by the player.

Mafia is also noted for having comprehensive damage physics on nearly all vehicles. While substantially more robust than their real counterparts, smaller and weaker vehicles stand less abuse before breaking down and finally exploding, than large armoured vehicles. More realism is added here compared to other games in the same genre, such as the ability to puncture the fuel tank, overheat the engine, and the ability to break transmission gears. Many exterior components (such as windows, tires, headlights, and bumpers) can be removed from most vehicles with physical means such as crash-driving, hitting with blunt weapons (fists, baseball bat) as well as firing weapons at them.

Mafia is set in the 1930s, between the fall of 1930 through to the end of 1938, during the later part of Prohibition, which ended in 1933. The game is set in the fictional American city of 'Lost Heaven' (loosely based on New York City, Chicago, and parts of New Jersey and Boston of the same time period).

Although the plot is not directly taken from the movie, certain names such as Paulie, and certainly the faces of the characters, the style and pace of the story, and many scene elements appear to be heavily inspired by the 1972 Francis Ford Coppola film "The Godfather," the 1984 Sergio Leone epic "Once Upon a Time in America" and the 1990 Martin Scorsese film "Goodfellas". Indeed, the front of the box playfully announces "Welcome to the game of the greatest 1930s gangster film Scorsese never made."

The player takes the role of taxi driver Thomas (Tommy) Angelo, who, while trying to make a living on the streets of Lost Heaven, unexpectedly and unwillingly becomes involved inorganized crime as a driver for the Salieri crime family, led by Don Salieri.

Through the events of the game's story, Tommy begins to rise through the ranks of the Salieri 'family', which is currently battling the competing Morello family, led by the sharply-dressed Don Morello. Eventually becoming disillusioned by his life of crime and violence, Tommy arranges to meet a detective (Detective Norman) in order to tell him his story, to be given witness-protection, and to aid the detective in the destruction of the Salieri crime family. The 'Intermezzo' chapters of the game depict Tommy sitting in a cafe with the detective, relating his life story and giving out important pieces of information at the same time.